28. Which of the following characteristics are expected in the first animals to have colonized land? 1. were probably herbivores (ate photosynthesizers); 2. had four appendages; 3. had the ability to resist dehydration; 4. had lobe-finned fishes as ancestors; 5. were invertebrates (A) 1, 3, and 5 (B) 1, 2, 3, and 4 (C) 3 only (D) 3 and 5